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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Heather and moorland view, near Birch tor, Dartmoor NP, Devon, UK, August 2008
Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK. January
Hartland Quay, rugged north Devon coastline, high tide at sunset, Devon, UK, April 2012
Star trails above Scots pine (Pinus sylvestris) woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, March 2017
Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Firefly squid (Watasenia scintillans) emitting light from photophores, Toyama Bay, Japan. This form of bioluminescence is used as counter illumination camouflage
Fishing cat (Prionailurus viverrinus) walking with fish in mouth. Andhra Pradesh, India. Camera trap image. Photo© Anjani Kumar/Felis Images
Pine marten (Martes martes) foraging in Pine (Pinus sp) woodland at night. Glenfeshie, Cairngorms National Park, Scotland, UK
Badger (Meles meles) emerging through gateway in wall at night. In urban area, Sheffield, England, UK. October 2018
RF - Maidens veil / Bridal veil fungus (Phallus indusiatus) with indusium fully formed, on rainforest floor. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Red fox (Vulpes vulpes) visiting woodland stream to drink at night. Camera trap image. Near Bath, England, UK. June
Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night
Vik i Myrdal church in winter, with full moon, Iceland. January 2015
Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Maned wolf (Chrysocyon brachyurus) searching for food, Piaui, Cerrado, Brazil, South America
Ice stalagmites / ice spikes formation, Lake Baikal, Siberia, Russia, March
Under water in the Abismo Anhumas or Anhumas Abyss. This is a 80 metre deep lake, at the bottom of a 72 metre deep cave. Bonito area, Serra da Bodoquena (Bodoquena Mountain Range)
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
Olm (Proteus anguinus) close up of front limb, these only have three toes whilst the rear limbs have only two. Captive, Slovenia
RF- Giraffe silhouette at sunset. Namibia, Etosha National Park
Black panther / melanistic Leopard (Panthera pardus) baring teeth, captive
Beech woodland in dense fog. Derbyshire, UK, November
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Mountains in snow, Aiguille du Tour with Tower Glacier in front. Sunset with a dark sky. Chamonix, Haute Savoie, France, December 2011
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Dusky leaf-nosed bat (Hipposideros ater) flying out of daytime cave roost in a sandstone escarpment, Kimberley, Western Australia
Oval squid (Sepioteuthis lessoniana) photographed at night, Green Island, Taiwan. Minimum fees apply
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic volumetric lighting. Vertes Mountains, Hungary
Andalusian stallion running in dark arena. Quebec, Canada
Young male Leopard (Panthera pardus) playing with kill after successfully hunting a young Impala at night. South Luangwa National Park, Zambia
African penguin (Spheniscus demersus) standing on rock at dusk, with clustered penguins in background. South Africa. November
European mantis (Mantis religiosa) hunting swarming Pale burrower mayfly (Ephoron virgo), mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August
White rhino (Ceratotherium simum) female and calf looking in opposite directions, reflected in waterhole at night. Zimanga private game reserve, KwaZulu-Natal, South Africa
Eurasian otter (Lutra lutra) feeding whilst on log in woodland burn, Glenfeshie, Cairngorms National Park, Scotland, UK, September
RF - Hartland Quay, late evening light, Hartland, Devon, UK. July
Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica
Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October
Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean
Blue shark (Prionace glauca) seen from above with pilot fish, Azores Islands, Portugal, Atlantic Ocean
Fly agaric (Amanita muscaria) in foggy forest, Los Alcornocales Natural Park, southern Spain, January
Fireflies {Lampyridae} in rainforest, Singapore
Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April
Snow goose (Anser caerulescens) two during migration with dark clouds against the sun, Quebec, Canada, October
Arctic Fox (Vulpes lagopus semenovi), in dark summer pelage, patrolling cliffside rocks along the coast of Medny Island in the Commander Islands (aka Komandorsky Islands), Russia, July
