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Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea
Yellow bellied snapping turtle (Elseya flaviventralis) about to surface for air, Arnhem Escarpment Creek, Northern Territory, Australia. June
Bluebell (Hyacinthoides non-scripta) agianst white background, Mull, Scotland
Liquidambar / Sweetgum (Liquidambar styraciflua) tree leaf changing to autumn colour, France
Borage (Borago officinalis) in flower with white background, Brechin, Scotland, UK, June
Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
Gyre flyingfish (Prognichthys glaphyrae) photographed in the Sargasso Sea, Atlantic Ocean
Jellyfish, probably a purple stinger (Pelagia noctiluca) at night in the Sargasso Sea, Atlantic Ocean
Winter aconite (Eranthis hyemalis) close up with white background
Great horned owl (Bubo virginianus) in flight. Saskatchewan, Canada, August
Columbus crab (Planes major) which was stranded on the beach, South Africa
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
California condor (Gymnogyps californianus) in flight, wings radio tagged. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. 2011
Seep monkey flower (Mimulus guttatus), bifid stigma above stamens. Nectar spot guides and central groove leading pollinators to nectar. Focus stacked
Snowdrop (Galanthus nivalis) anthers and inner petals. Outer petals have been removedSnowdrop (Galanthus nivalis) anthers and inner petals.Outer petals have been removed. Focus stacked
Himalayan balsam (Impatiens glandulifera), dissection of flowers. Male phase above, downward projecting stamens release pollen onto backs of bumblebees and honey bees
Sweet breath of spring (Lonicera fragrantissima) in garden, Surrey, England, UK. Native to China
Scarlet monkey flower (Mimulus cardinalis). The high position of the stamens and stigma ensure that the heads of hummingbird pollinators make contact with them whilst nectaring
Gold band lily (Lilium auratum) flower. Yellow nectar guides lead pollinators to nectaries. Pollinators are swallowtail butterflies and hawkmoths. Cultivated in garden, Surrey, England, UK
Cherry laurel (Prunus laurocerasus) flowers. England, UK. April. Controlled conditions
White dead nettle (Lamium album) with stamens protected from rain by overhanging upper lip. Pollinated by bumblebees. Naturalised in garden, Surrey, England, UK. May
Bird of paradise / Crane flower (Strelitzia reginae). Cultivated in garden. Native to South Africa
Glory lily (Gloriosa superba) bud and flower with reflexed petals and trifid stigma. Pollinated by butterflies. National flower of Zimbabwe
Mandrinette (Hibiscus fragilis), cultivated in breeding program at Kew Gardens, London, UK. Endemic to Mauritius
Snowdrop (Galanthus nivalis) dispersing pollen by sonication which replicates buzz pollination. Controlled conditions
Chilean bellflower (Crinodendron hookerianum) releasing pollen via sonication which replicates buzz pollination. Native to Chile. Controlled conditions
New Zealand tree fuchsia (Fuchsia excorticata) with blue pollen and yellow tipped stigma. Focus stacked
Bloody bellflower (Nesocodon mauritianus), female phase with red nectar surrounding stigma. Native to Mauritius. Cultivated at Kew Gardens, London, England, UK
Cup and saucer plant (Cobaea scandens), stamens retracting and a drop of nectar on petal tip. Cultivated in conservatory, Surrey, England, UK. Controlled conditions, focus stacked
Goat willow (Salix caprea) female catkin in UV light, fine hairs and nectar fluroescing. Surrey, England, UK. Focus stacked
Hedgehog-shaped lip bulbophyllum (Bulbophyllum echinolabium), Cultivated in glasshouse, Kew Gardens, London, England, UK. Native to Sulawesi, Indonesia. Controlled conditions
Gold band lily (Lilium auratum) with yellow nectar guides. In cultivation. Native to Japan
Morning glory (Ipomoea tricolor) flower with pollen grains scattered by visiting insects
Southern magnolia / Bull bay (Magnolia grandiflora) flower. The opening and closing of petals controls the time beetle pollinators enter and leave the flower
Musquia (Musschia aurea) flower. Cultivated in glasshouse, UK. Pollinated by Lizard (Lacerta dugesii) and Madeira bumblebee (Bombus maderensis) in Madeira
Winter aconite (Eranthis hyemalis) with ring of nectaries below stamens. Focus stacked
Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK
Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2
Porcelain orchid (Chloraea magellanica) with osmophores on lip fluorescing in UV light. Chile. Controlled conditions, focus stacked. Sequence 2/2
Common tern (Sterna hirundo) in flight, Norfolk, UK, August
Grey heron (Ardea cinerea) head profile, Regents Park, London, UK August
Common / Bandit pipistrelle bat (Pipistrellus pipistrellus) hunting at twilight, Berwickshire, Scotland, August
Violet-green Swallow (Tachycineta thalassina), female in flight carrying a feather for nest lining, spring, Mono Lake, California, USA, July
European Otter (Lutra lutra) standing on ice. Captive. The Netherlands, January
European Otter (Lutra lutra) running over snow. The Netherlands, December. Captive
Two tabby kittens, Smudge and Picasso, age 9 weeks
RF- Head portrait of Bulldog puppy with chin on paws