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African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa
Two Arctic hares (Lepus arcticus) fighting, Northeast Greenland National Park, Greenland. Veolia Environnement Wildlife Photographer of the Year 2009: Mammal Behaviour category: Runner-up
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Mountain Hares (Lepus timidus) boxing in winter. Cairngorms National Park, Scotland, UK, January
Wild Horse / Mustang shaking head and mane, Adobe Town Herd Area, southwestern Wyoming, USA, WINNER OF WILD HORSE SECTION, EQUINE IDEAL PHOTOGRAPHY CONTEST 2012
Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK
White tailed sea eagle (Haliaeetus albicilla) interacting with Great egret (Ardea alba) Lake Csaj, Kiskunsagi National Park, Hungary. January
Great egret (Ardea alba) and Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname
Kottigehara dancing frog (Micrixalus kottigeharensis), dancing frog name given because of foot flagging behavior between competing males. Endemic to Western Ghats. Agumbe, Karnataka, India
Common / Eurasian cranes (Grus grus) three 18 month juveniles, Bart, Christine and Gilbert, released by the Great Crane Project onto the Somerset Levels and Moors
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition
Two European goldfinches (Carduelis carduelis) fighting over food in snow, Wrocław, Lower Silesia, Poland. February
Two Ruffs (Philomachus pugnax) male, competing at a lek, Vardo, Norway. May
Black grouse (Tetrao tetrix) males fighting in snow during the breeding season, Viiksimo, Finland. April
Three Spotted hyenas (Crocuta crocuta) fighting over meat from a nearby carcass with flock of White-backed vultures (Gyps africanus) watching in background, Mana Pools National Park, Zimbabwe
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Dalmatian pelican (Pelicanus crispus) group squabbling over fish, Lake Kerkini, Greece
Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. February
White tailed eagles (Haliaeetus albicilla) two juveniles fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Great egret (Ardea alba) fighting over fish with Grey heron (Ardea cinerea) Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Ruffs displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Ruff displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Two Lionesses (Panthera leo) trying to defend kill from large pack of Spotted hyaenas (Crocuta crocuta) Masai-Mara game reserve, Kenya
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
A wild Misaki-uma breeding stallion (right) brings back one of his mares to his band while another breeding stallion (left) charges him away, in the Cape Toi Reserve, Miyazaki Prefecture
