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Grey seal (Halichoerus grypus), male and female, mating behaviour, Heligoland, Germany
Iceberg and reflection, in Rode Fjord (Red Fjord), Scoresby Sund, Greenland, August
Icebergs and autumn tundra near Rode O (Red Island) in Rode Fjord (Red Fjord), Scoresby Sund, Greenland, August
Iceberg and hills in Rode Fjord (Red Fjord), Scoresby Sund, Greenland, August
Orca (Orcinus orca) hunting South American sealion (Otaria flavescens) close to shore. Punta Norte, Valdez Peninsula, Argentina. April
Traigh Scarasta on the Isle of Harris. Isle of Harris, Outer Hebrides, Scotland, UK, August
Weddell seal (Leptonychotes weddellii) hauled out on ice, Antarctic Peninsula, Antarctica
Bull Skink (Liopholis multiscutata) in coastal dunes at Browns Beach on Kangaroo Island, South Australia, Autumn
Aerial view of a blue hole on Eleuthera Island, Bahamas
Feral horse (Equus caballus) walking on a remote sandy beach, near Arta, Mallorca, Spain, August
Lapwing (Vanellus vanellus) Titchwell RSPB Reserve, Norfolk, England, UK. November
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Marine iguana (Amblyrhynchus cristatus), Punta Vicente Roca, Isabela Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos
Golden cownose rays (Rhinoptera steindachnyri) schooling in sheltered mangrove lagoon, Academy Bay, Santa Cruz Island, Galapagos, Ecuador
Sequence of strips of cultivated Machair with early shoots of Black oats (Avena strigosa) pushing through the sand. North Uist, Scotland, UK, June
King penguins (Aptenodytes patagonicus) walking in line on a windy beach. Sanders Island, Falklands. November
Aerial view of Carters Beach on the Southshore of Nova Scotia, Canada. August
Lowland blotched blue-tongue (Tiliqua nigrolutea) on a coastal bluff in the Bay of Fires region of north-eastern Tasmania, Australia, spring
Lemon-yellow tree frog (Hyla savignyi) peering around coastal plant. Cyprus. April
Seaweeds including Thongweed / Sea thong (Himanthalia elongata), Tangleweed kelp (Laminaria digitata), Toothed wrack (Fucus serratus) and Dulse (Palmaria palmata)
Rock pipit (Anthus petrosus) foraging for invertebrates among seaweed on the strand line as the tide rises at sunset, Cornwall, UK, September
Otter (Lutra lutra) portrait on shore with boats in background. Argyll, Scotland, UK, August
Otter (Lutra lutra) feeding, Argyll, Scotland, UK, August
Yellow horned poppy (Glaucium flavum) growing on shingle beach, Norfolk, UK, August
Dog walkers walking along The Wash in evening low tide, Norfolk, UK August 2013
Chocolate working cocker spaniel amongst sand dunes. Formby, Merseyside, UK
Sea stacks at Mangurstadh / Mangersta Beach, Isle of Lewis and Harris, The Outer HebridesSea stacks at Mangurstadh/Mangersta Beach, Isle of Lewis and Harris, The Outer Hebrides, Scotland, UK. October
Luskentye beach, mountains and incoming tide, Isle of Lewis and Harris, Outer Hebrides, Scotland, UK. October 2018
Bamburgh Castle and sand dunes, late evening light, Bamburgh, Northumberland, UK. March 2018
Godrevy Lighthouse at dusk, St. Ives Bay, West Cornwall, England, UK. February 2016
RF - Hartland Quay, late evening light, Hartland, Devon, UK. July
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Feral sheep with Galtachan islands behind, Shiant Isles, Outer Hebrides, Scotland, UK. June 2018
Razorbill (Alca torda) orange sea lichen (Caloplaca marina) covered roch behind, Shiant Isles, Outer Hebrides, Scotland, UK. June
Mangroves on Tupeti Island, Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Rock flagtail (Kuhlia rupestris) fish in river on the Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Griffon vulture (Gyps fulvus) landing on coast next to flock. Cantabria, Spain, March
Greater flamingos (Phoenicopterus roseus) flock, Sado Estuary, Portugal. January
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Lemon shark pups (Negaprion brevirostris) spend the first 5-8 years of their life in mangrove forests. The tangle of roots provides protection from predators
Part of a submerged Holocene forest, revealed after a winter storm from beneath the sand at Hells Mouth (Porth Neigwl), Abersoch Wales, UK March 2014
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Killer whale (Orcinus orca) hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Common eider (Somateria mollissima), female on her nest with sea behind, Finland, May
