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Violet bramble rust (Phragmidium violaceum) causing red colouration on Bramble leaf (Rubus fruticosus aggregate). Peak District National Park, Derbyshire, UK. October
Raft Spider (Dolomedes fimbriatus) female on heathland pool. Note wing of predated damselfly floating on waters surface under the spider. Dorset, UK. August
Sand lizard (Lacerta agilis) male basking. Dorset, England, UK, May
Sanderling (Calidris alba) running along the tide-line, reflected in the mud, Norfolk, England, UK. December
Lavender (Lavandula angustifolia) fields, Valensole Plateau, Alpes Haute Provence, France, June
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Fallen leaves in pond with aquatic plants, Denmark, Europe, September
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Galapagos green turtle (Chelonia agassizii) surfaces to breathe. Floreana Island, Galapagos Islands, Ecuador. East Pacific Ocean
Ancient dead Camelthorn tree (Vachellia erioloba) trees with red dunes and mist, Namib desert, Deadvlei, Sossusvlei, Namibia. August 2015
Avenue of Oak trees (Quercus) Eickelberg, Warnowtal, Germany, September
Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India
Mountain landscape, Svalbard, Norway, July
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
View of Stokksnes Peninsula, seen from Hofn, Iceland, March 2014
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Lioness (Panthera leo) and cubs in rain, Masai Mara Game Reserve, Kenya, September
Sunset off Scott Island, with floating pieces of sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August 2015
Marbled white butterfly (Melanargia galathea) resting on grass head, backlit against black background, Dunsdon, Devon, UK. July 2014
Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa
Bearded seal (Erignathus barbatus) hauled out on ice, Spitsbergen, Svalbard, Norway, September
Great white egret (Egretta alba or Ardea alba) catching cyprionid fish or carp in fish farm, Pusztaszer Protected Landscape, Kiskunsagi, Hungary, May
Lion (Panthera leo) male resting, Masai Mara Game Reserve, Kenya, November
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Coastal brown bear (Ursus arctos) fishing, Lake Clarke National Park, Alaska, September
Porcupine (Erethizon dorsatum) on tundra. North Slope, Alaska, USA. September
Mallard duck (Anas platyrhynchos) drake swimming through reflection of house, Bude canal, Cornwall, UK. April 2015
Common darter dragonflies (Sympetrum striolatum) two females resting on reed, Dunsdon Nature Reserve, Devon, UK. August
Barred owl (Strix varia) stretching wings, Corkscrew Swamp Audubon Sanctuary, Florida, USA, March
Waterfalls cascading between mountain lakes, Plitvice Lakes National Park, Croatia. January 2015
Porcelain fungus (Oudemansiella mucida) gills. Peak District National Park, Derbyshire, UK. October
Greenfinch (Carduelis chloris) pair, one perched on branch and one hovering in snowfall, Derbyshire, February
Nudibranch (Flabellina lineata) in aquarium, taken in the field in Gulen, Norway. North East Atlantic Ocean. Collected for scientific research
Little egret (Egretta garzetta) shaking water off wings at sunset, Lake Csaj, Pusztaszer, Hungary. May. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Polar bear (Ursus maritimus) mother and large second year cub swim across calm water, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Green turtle (Chelonia mydas) resting in the shallows of the coast, Bijagos Islands, Guinea Bissau. Endangered species
Aerial view of Amazon Rainforest, and the Manati River, Peru, July 2015
Firewheel (Gaillardia pulchella) flowers at twilight, Laredo Borderlands, Texas, USA. April
