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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Red squirrel (Sciurus vulgaris), two curled up asleep in drey inside nest box. Nest of lichen and Pine needles. Highlands, Scotland, UK. Medium repro only
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Snowdrop (Galanthus Sp. ) single flower in snow, Buckinghamshire, England, UK, FebruarySnowdrop (Galanthus Sp.) single flower in snow, Buckinghamshire, England, UK, February
Eastern tiger salamander (Ambystoma tigrinum) North Florida, USA. December
Canadian timber wolf / Northwestern wolf (Canis lupus occidentalis) pack moving through snow, Wood Buffalo National Park, Alberta, Canada. Taken on location for BBC Frozen Planet, March 2009
House Martin (Delichon urbica) Spain, April
Tre Cime di Lavaredo / Drei Zinnen, three distinctive mountain peaks in the Sexten Dolomites seen from WW1 cave shelter, South Tyrol, Italy, October 2019
English bluebell (Hyacinthoides non-scripta) Lower Woods, Gloucestershire, England, UK, April
Elephant (Loxodonta africana) herd walking in a line, Etosha National Park, Namibia
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Buckholt Wood is a beech (Fagus sylvatica) wood and part of the Cotwolds Commons and Beechwoods National Nature Reserve, on the scarp slope of the Cotswolds near Birdlip, Gloucestershire
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March
Golden eagle (Aquila chrysaetus), Kuusamo, Finland, January
Grey wolf running in snow (Canis lupus), Minnesota, USA. January. Controlled situation
Lady Amhersts pheasant (Chrysolophus amherstiae) Kanding, Sichuan, China
Golden eagle (Aquila chrysaetos) Sodankyla, Lokka, Finland, April
Sea elephant (Cardiopoda placenta), marine pelagic gastropod mollusc in the family Pterotracheidae. Deep sea heteropod. Atlantic Ocean, close to Cape Verde. Controlled conditions
Deadly nightshade (Atropa belladonna) Quarry Hangers (SWT), Chaldon, Surrey, England, June
Red fox (Vulpes vulpes) cubs sleeping curled up together, Switzerland
Central bearded dragon (Pogona vitticeps) male on stick in chenopod shrubland habitat, Meringur in NW Victoria, Australia
Autumn quaking aspen trees (Populus tremuloides) below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
White-tailed deer (Odocoileus virginianus) doe standing in snow, New York, USA
Common octopus (Octopus vulgaris) on a coral reef, The Bahamas. August
Golden snub-nosed monkey (Rhinopithecus roxellana), mother and baby, Qinling Mountains, Shaanxi province, China
Face portrait of a Great hammerhead shark (Sphyrna mokarran) off Bimini, Bahamas
Eclectus parrot (Eclectus roratus) two males arrive at a nest cavity at the same time, both to provision the female within. Iron Range National Park, Cape York Peninsula, Queensland, Australia
Pallass cat (Otocolobus manul) licking lips whilst running over ice. East Mongolia. February
Aerial view of Glassy Point, East Coast of Dominica, West Indies. December 2019. Drone photo. Stitched panorama
Hyacinth macaws (Anodorhynchus hyacinthinus) squabbling on palm tree, Pantanal, Mato Grosso, Brazil
Lion (Panthera leo) males mock fighting / play fighting. Ngorongoro Crater, Ngorongoro Conservation Area (NCA), Tanzania
Red fox (Vulpes vulpes) running through snow. Hayden Valley, Yellowstone, USA. January
Fosa / Fossa (Cryptoprocta ferox) male resting in dry decidous forest. Kirindy, western Madagascar. Endangered
Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina
Common Loons (Gavia immer) one diving underwater, Michigan, USA. June
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Dramatic sunset with storm clouds over Roseau, Caribbean sea view in Dominica, Lesser Antiles. September 2019
Great Grey Owl (Strix nebulosa) Kuhmo Finland, March
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Golden jackal (Canis aureus) pup sitting on a grassy verge by a path, Danube Delta, Romania, July
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Common Cuckoo (Cuculus canorus) perched on gate Surrey, England, UK. April
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
