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Grey heron (Ardea cinerea) in rain with another heron behind, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Globe Thistle (Echinops ritro), dead in winter landscape with Alpine ibex (Capra ibex) silhouet behind, Gran Paradiso national park, Italian Alps, Italy, January
Katydid (Tettigoniidae) with Giraffe (Giraffa camelopardalis) in the background, Masai Mara National Park, Kenya
Gannet (Morus bassanus) nearest out of focus, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Common blue butterflies (Polyommatus icarus) soft focus resting on grass, Vealand Farm Devon Wildlife Trust Reserve, Devon, England, UK. September
Great egret (Ardea alba) group of five at waters edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February
RF - Eschers blue butterfly (Polyommatus escheri) on grass, with soft focus blue flowers, Aosta Valley, Gran Paradiso National Park, Italy
Hoopoe (Upupa epops) with European roller (Coracias garrulus) in the background. Pusztaszer, Hungary. May
Dew on spiderweb with Common poppy (Papaver rhoeas) in the background, Sierra de Grazalema Natural Park, El Bosque, southern Spain, April
Dew on spiderweb with Grey-leaved cistus (Cistus albidus) Sierra de Grazalema Natural Park, El Bosque, southern Spain, April
Orchid (Orchis langei) in sunset light, Sierra de Grazalema Natural Park, southern Spain, May
Orchid (Orchis olbiensis), Sierra de Grazalema Natural Park, Benaocaz, southern Spain, May
Lion (Panthera leo) portrait of two lionesses, captive, occur in Africa
African lioness (Panthera leo) sitting patiently in the long grass, watching herd of Common eland (Tragelaphus oryx). Focus on lion. Grumeti Reserve, Northern Tanzania
Common blue damselfly (Enallagma cyathigerum) roosting among dewy grasses, Tamar Lakes, Cornwall, UK. June
RF - Common blue butterflies (Polyommatus icarus) resting on a backlit grass, evening light, Vealand Farm, Devon, UK. June
RF - Wild garlic / Ramson flowerts (Allium ursinum) Valency Valley, Boscastle, Cornwall, UKRF - Wild garlic/Ramson flowerts (Allium ursinum) Valency Valley, Boscastle, Cornwall, UK. May
Poppy (Papaver rhoeas) Groot Schietveld, Wuustwezel, Belgium, July
Demoiselle cranes (Anthropoides virgo) flock, India
RF- Mute swan (Cygnus olor) close up peering at camera, juvenile. Fife, Scotland, UK, November
Tropical Screech Owl (Megascops choliba) perched in tree hole in rainforest canopy. Cristalino State Park, Amazonia, Brazil, South America
Lesser spotted fritillary (Melitaea trivia) adult at rest on yellow flower, Bulgaria
Chestnut-headed Tesia (Tesia castaneocoronata) perched, Sichuan Province, China
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Great White Egret (Egretta alba) with water splashing from beak, Hungary January
Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Marbled white butterfly (Melanargia galathea) resting on reed, Devon, UK, July
Cowslips (Primula veris) growing on organic farm. Norfolk, UK, April
Two male Bearded reedling (Panurus biarmicus) eating seeds from the spike of a common bulrush (Typha latifolia) in a reedbed, Belgium, January. Digital composite
Small Yellow Underwing moth (Panemeria tenebrata) on stitchwort (Stellaria) South Karelia, southern Finland, June
Northern Hawk-Owl (Surnia ulula) seen through branches, southwest Finland, February
Scottish wildcats (Felis sylvestris), captive, UK, June
Spectacled caiman (Caiman crocodilus) resting in shrinking pool, Pouso Alegre lodge. Pantanal, Brazil. Highly commended in the Other Animals Category of the GDT competition 2013
Kittens exploring garden lawn, Germany
Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Natures Best Photography Windland Smith Rice International Awards Competition
Golden-crowned Sifaka or Tattersalls Sifaka (Propithecus tattersalli) climbing down tree in forests near Andranotsimaty, Daraina, north east Madagascar. Critically Endangered
Phayers Leaf monkey (Trachypithecus phayrei) Gaoligongshan NP, Yunnan province, China
Greater Stitchwort (Stellaria holostea) and Bluebells (Hyacinthoides non-scripta) in flower, UK, May
Swallowtail (Papilio machaon) in flight, Finland, Europe
Female goshawk (Accipiter gentilis) in flight, just after taking off from perch. Southern Norway, January
Fallow deer (Dama dama) one deer stops and looks whilst the herd runs on, with sunlit leaves out of focus, Vastmanland, Sweden
Pair of Emerald damselflies (Lestes sponsa) resting on a reed, with only tips of wings in focus, Cornwall, England, UK, August
RF - Common spotted orchids (Dactylorhiza fuchsii), Volehouse nature reserve, Devon, UK, June
Wild garlic / Ramsons (Allium ursinum) flowering in woodland, Lanhydrock Woodland, Cornwall, UK, May
RF- Beech (Fagus) and Bluebell (Hyacinthoides non-scripta) woodland at Lanhydrock, Cornwall. UK
Female Jaguar (Panthera onca), captive, occurs in Southern and Central America
Razorbill (Alca torda) pair calling, between soft focus vegetation. Great Saltee, Saltee Islands, Co. Wexford, Ireland, June
White stork (Ciconia ciconia) adult portrait, captive, Vogelpark Marlow, Germany, May