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Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Common blue butterfly (Polyommatus icarus) resting on Devil's bit scabious (Succisa pratensis), Volehouse Farm, Devon, UK. August
Leopard (Panthera pardus) cub looking up at birds (out of frame) with mother in background, Khwai River, Botswana
Fiery throated hummingbird (Panterpe insignis) an endemic bird species. Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Skewbald and chestnut Icelandic horses in the snow, Snaefellsnes Peninsula, Iceland, March
Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
RF- Head portrait of a European grey wolf (Canis lupus), with a second in the background, captive, Norway, February
RF - Red Deer stags (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Red fox (Vulpes vulpes) peering from behind tree with bluebells in foreground, Cheshire, June
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
Fireweed / Rosebay willowherb (Chamerion angustifolium), Klein Schietveld, Brasschaat, Belgium, July
RF - Marbled white butterfly (Melanargia galathea) on knapweed, with soft focus leaves, Aosta Valley, Gran Paradiso National Park, Italy
Leaves emerging on European beech tree (Fagus sylvatica) Angus, Scotland, UK. May
Wild Garlic / ramsons (Allium ursinum) leaves emerging in early spring. Peak District National Park, Derbyshire, UK. March
Bottlenose dolphins (Tursiops truncatus) approaching with curiosity, Dolphin Reef, Eilat, Israel. Red Sea
Common poppies (Papaver rhoeas) backlit in evening light, Polly / Porth Joke, Pentire, Newquay, Cornwall, UK, July
Luskentye beach, mountains and incoming tide, Isle of Lewis and Harris, Outer Hebrides, Scotland, UK. October 2018
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Yellow wagtail (Motacilla flava) perched among flowering Rapeseed (Brassica napus), Scotland, UK. May
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
Darjeeling bush frog (Raorchestes Annandalii) croaking, sitting on a petal, Kalimpong, West Bengal, India
Crab spider (Thomisus onustus) on white flowerhead, Sierra de Grazalema Natural Park, southern Spain, May
Spoonwing / Thread-winged antlion (Nemoptera bipennis) on flower, Sierra de Grazalema Natural Park, southern Spain. June
Red Campion (Silene dioica), backlit in early morning sunlight, spring, Broxwater, Cornwall, UK. April
RF - Common blue butterflies (Polyommatus icarus) resting on grass. Vealand Farm, Devon, UK. September
Rock sea-spurrey (Spergularia rupicola) growing on seashore at the base of a cliff, Cornwall, UK, September
Small tortoiseshell butterfly (Aglais urticae) resting on heather, Westhay, Somerset Levels, UK, August
Satiny Parrot snake (Leptophis depressirostris) Santa Rita, Costa Rica
Hawthorn berries (Crataegus monogyna) in Poplar woodland (Populus) Sierra de Grazalema Natural Park, southern Spain. October
Head portrait of grey almost white Andalusian mare running in snow, Berthoud, Colorado, USA. January
RF- Capercaillie (Tetrao urogallus) male displaying, Inshriach, Cairngorms National Park, Scotland, UK. February
Grasshopper Warbler (Locustella naevia) singing, Joensuu, Finland, May
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Common poppy (Papaver rhoeas) bud, Polly / Porth Joke, Pentire, Newquay, Cornwall, UK, July
Marsh harrier (Circus aeruginosus) adult female circling over Oilseed rape crop, Norfolk, England, May. Digital composite
Redpoll (Carduelis flammea) male landing on Alder (Alnus glutinosa) Surrey, England. February
Eyelash palm pit viper (Bothriechis schlegelii) with tongue extended, Mindo, Pichincha, Ecuador
Clarion Burrowing Owl (Athene cunicularia rostrata), Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Alpine meadow in flower including Scabious flowers. Austrian Alps at 1700 metres altitude, North Tyrol, Austria, June
RF- Common frog (Rana temporaria) portrait, Cornwall, UK, January
RF - Early Purple Orchid (Orchis mascula) in flower, Broxwater, Cornwall, UK. May 2016
RF- Daffodils (Narcissus) in flower photographed using soft focus technique
Indochinese tiger (Panthera tigris corbetti) looking up, behind screen of leaves, captive, occurs in South East Asia
RF- Pool Frog (Rana lessonae) sitting on white lily pad, Danube delta rewilding area, Romania
