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Harpy Eagle (Harpia harpyja) portrait. Gamboa, Soberania National Park, Panama
Dipper (Cinclus cinclus) singing from prominent rock in breeding season, Derbyshire, UK
Blacktail Prairie Dog (Cynomys ludovicianus) engaging in Jump-yip behavior - A strong arch of the back or " jump" followed by a shrill " yip" - Thought to occur when a predator
Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) screaming and showing its teeth, Foping Nature Reserve, Shaanxi, China. Endangered species
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Raven (Corvus corax) calling in the snow, Kemijarvi, Finland, February
Red deer (Cervus elaphus) male stag roaring portrait, rut season, Jura, Scotland, UK, September
Red deer stag (Cervus elaphus) calling during rut. Richmond Park, England, UK, Europe
Male Mantled howler monkey (Alouatta palliata) on foot bridge, La Selva Biological Station, Costa Rica
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Javan leopard (Panthera pardus melas) roaring, native to the Indonesian island of Java. Captive. Digital composite
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Black spider monkey (Ateles chamek) calling, captive, Peru
Red howler monkey (Alouatta seniculus) howling, captive, digitally modified
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June
Capercaillie (Tetrao urogallus) adult male displaying in pine forest. Cairngorms National Park, Scotland, February
Tree swallows (Tachycineta bicolor), perched on wire, calling aggressively to each other, Madison River, Montana, USA, June
Two European grey wolves (Canis lupus) howling in winter landscape, captive. Bayerischerwald National Park, Germany
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Red deer (Cervus elaphus) stag bellowing during rut, at night with full moon, Derbyshire, UK
Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016
Wood Warbler (Phylloscopus sibilatrix) singing from perch. Wales, May
Red deer (Cervus elaphus) stag roaring at dusk during rutting season, Cheshire, UK October
RF - Sea otter (Enhydra lutris) floating, California, USA, July
Domestic goat (Capra hircus) pygmy goat bleeting, portrait, UK
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Common Crane (Grus grus) displaying in snow, Kuusamo Finland, April (Some background added.)
Darjeeling bush frog (Raorchestes Annandalii) croaking, sitting on a petal, Kalimpong, West Bengal, India
Common tern (Sterna hirundo) flying low over water. Hjalstaviken, Uppland, Sweden
Fallow deer (Dama dama) buck bellowing in autumn vegetation, Bradgate Park, Leicestershire, UK, October
Red deer (Cervus elaphus) stag roaring during rut, with resting females, England, UK. September
Very young tabby kitten, 6 weeks, with raised paw and open mouth
Corncrake (Crex Crex) calling, with breath vapor in early morning, Kiskunsagi National Park, Hungary
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Black korhaan (Eupodotis afra) male displaying and calling. Etosha National Park, Namibia, March
Red deer (Cervus elaphus) stag bellowing, Los Alcornocales Natural Park, southern Spain, September
Arctic fox (Vulpes lagopus) in summer coat calling, Wrangel Island, Far Eastern Russia, August
Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India
Anils bush frog (Raorchestes anili) inflating vocal sac to attract mate, Endemic to Western Ghats. Coorg, Karnataka, India
Jayarami Bush frog (Raorchestes jayarami), inflating vocal sac, calling to attract mate, Endemic to Western Ghats. Anamalai Wildlife Sanctuary, India
Wrybill (Anarhynchus frontalis) standing shallow water with beak wide open. Lake Ellesmere, South Island, New Zealand. November
RF - Bald eagle (Haliaeetus leucocephalus) calling, captive, occurs in North America
