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Black blotched pufferfish (Diodon liturosus) inflated with water at night as defensive behaviour, Sipadan Island, Borneo, Malaysia
Rafflesia flower (Rafflesia keithii) in Gunung Gading NP, Borneo, Sarawak, Malaysia
Orang utan baby (Pongo pygmaeus) head portrait, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Orang utan (Pongo pygmaeus) baby hanging from trees, and chewing on fingers, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Waglers / Temple pit viper (Tropidolaemus wagleri) in lowland rainforest, Gunung Palung National Park, Borneo, West Kalimantan, Indonesia
Head portrait of Bornean Horned Frog (Megophrys nasuta) among the leaf litter in the lowland rainforest of Borneo. Danum Valley Conservation Area, Sabah, Malaysia
Strangler Fig (Ficus sp. ) that has killed its host tree long ago. The host has rotted awayStrangler Fig (Ficus sp.) that has killed its host tree long ago. The host has rotted away, leaving a hollow center. Lowland rainforest in Borneo. Gunung Palung National Park, Indonesia
A waterfall in the lowland rainforest of Borneo, with pool surrounded by ferns and other plants. Lambir Hills National Park, Sarawak, Malaysia
Aerial view of rivers and mangrove forest. Sarawak Mangrove Reserve, Sarawak, Borneo, Malaysia. June 2006
Grey gibbon (Hylobates muelleri) swinging from branch in rainforest and using foot to grip plant vine, Mount Kinabalu NP, Sabah, Borneo, Malaysia
Red leaf monkey (Presbytis rubicunda) mother and baby, Danum Valley, Sabah, Borneo, Malaysia August
Dawn over the Segama River, with mist hanging over lowland rainforest. Heart of Danum Valley, Sabah, Borneo
Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Mist hanging over Lowland Dipterocarp Rainforest just after sunrise. Danum Valley Conservation Area, Sabah, Borneo
Two Millipedes (Julidae) close up portrait, Mulu, Sarawak, Malaysia
Great anglehead lizard (Gonocephalus grandis) resting on tree trunk, Mulu, Sarawak, Malaysia
Black-headed cat snake (Boiga nigriceps) coiled, portrait, Kubah National Park, Sarawak, Malaysia
Male Gold-specs jawfish (Opistognathus randalli), with mouth brooding eggs, the eyes can clearly be seen in the eggs indicating they will soon hatch, Mabul Island, Sabah, Malaysia, Celebes Sea
Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp)Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp.), Kubah National Park, Sarawak, Borneo
Close up of hands of a Bornean Orangutan (Pongo pygmaeus), Tanjung Puting National Park, Borneo
Orange backed red langur (Presbytis rubicunda chrysea) in rainforest understorey. Danum Valley, Sabah, Borneo, Malaysia
Infant Bornean Orang-Utan (Pongo pygmaeus) clinging to its mother. Camp Leakey, Tanjung Puting NP, Kalimantan, Borneo
Carrion fly on the parasitic plant Rafflesia pricei flower, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
Montane horned frog (Megophrys kobayashii) head close up, Kinabalu National Park, Sabah, Borneo, Malaysia
Minute / Dwarf filefish (Rudarius minutus) just over an inch long as an adult, Celebes Sea, Mabul Island, Malaysia
Sunda leopard cat (Prionailurus javanensis) kitten. Sabah, Malaysian Borneo
Blyths paradise flycatcher (Terpsiphone affinis) male calling, Sabah, Malaysian Borneo
Proboscis monkey (Nasalis larvatus) male leaps between trees. Sabah, Malaysian Borneo
Lowland dipterocarp rainforest, with huge Tualang (Tapang) or Mengaris Tree (Koompassia excelsa) in the foreground, Danum Valley, Sabah, Borneo
Western / Horsfields tarsier (Tarsius bancanus) hunting invertebrate prey in rainforest understory vegetation at night. Danum Valley, Sabah, Borneo
Giant shield bug nymphs (Pycanum sp. ), recently hatched from eggsGiant shield bug nymphs (Pycanum sp.), recently hatched from eggs. Danum Valley, Sabah, Borneo
Safety in Numbers - Giant Shield bug nymphs (Pycanum sp.), recently hatched from eggs, with a final individual still to emerge
Arboreal tarantula (Lampropelma sp. ) female guarding spiderlings at entrance to hollowArboreal tarantula (Lampropelma sp.) female guarding spiderlings at entrance to hollow retreat, 4 metres up a tree. Danum Valley, Sabah, Borneo
Lantern bug (Pyrops whiteheadi), detail of wing. Danum Valley, Sabah, Borneo
Lantern bug (Pyrops whiteheadi), detail of head. Danum Valley, Sabah, Borneo
Lowland dipterocarp rainforest at dawn, Danum Valley, Sabah, Borneo
Sumatran pitviper (Trimeresurus sumatranus), detail of scales. Danum Valley, Sabah, Borneo
RF - Maliau Falls, Maliau River. Centre of Maliau Basin - Sabah, Borneo. (This image may be licensed either as rights managed or royalty free.)
Portrait of Bornean Horned Frog (Megophrys nasuta) amongst leaf-litter in forest floor. Taken at night. Danum Valley, Sabah, Borneo
Crinoid (Crinoidea) on Soft coral (Alcyonacea). Derawan Islands, East Kalimantan, Indonesia
Pink anemonefish (Amphiprion perideraion). Derawan Islands, East Kalimantan, Indonesia
Flower of locally endemic Rafflesia (Rafflesia tengku-adlinii) flower diameter 22cm blooming on forest floor within lowland Dipterocarp rainforest, Maliau Basin, Sabahs Lost World, Borneo
Mist and low cloud hanging over lowland Dipterocarp Rainforest, just after sunrise, Danum Valley, Sabah, Borneo, May 2011
Close up of decaying leaf from rainforest floor, showing lichens and moulds. Danum Valley, Sabah, Borneo
One of the dams constructed by the local field staff of the Borneo Nature Foundation in the canals in the Sabangau (peat-swamp) Forest. These are made to combat forest fires
Pandan (Pandanus sp. ) growing in the Sabangau River, Central Kalimantan, IndonesiaPandan (Pandanus sp.) growing in the Sabangau River, Central Kalimantan, Indonesia
Maroon / Red leaf monkey / Langur (Presbytis rubicunda) eating fruits in tree, Danum Valley Conservation Area, Sabah, Borneo, Malaysia
Red leaf monkey (Presbytis rubicunda) female feeding in Strangler fig tree (Ficus dubia) Gunung Palung National Park, Kalimantan, Borneo, Indonesia