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Southern elephant seal (Mirounga leonina) beach master / bull roaring amongst females and pups, in breeding colony. St Andrews Bay, South Georgia. October
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Young grey seal pup (Halichoerus grypus) recently born on a beach in Orkney, Scotland, UK, April
Colonies of King penguins 1+Aptenodytes patagonicus+1 and Southern Elephant Seals 1+Mirounga leonina+1 on beach at St. Andrews Bay, South Georgia. November
Himalayan honey bees (Apis dorsata laboriosa) on comb, Mount Namjagbarwa, Yarlung Zangbo Grand Canyon National Park, Tibet, China. October
Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos
Northern gannet (Morus bassanus) in flight over colony, Quebec, Canada, August
Caribbean Flamingo (Phoenicopterus ruber) chick group walking around the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Aerial view of Caribbean Flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
King penguin (Aptenodytes patagonicus) colony. Right Whale Bay, South Georgia Island Panoramic composite view
A panoramic composite image of a massive grouping of King penguins (Aptenodytes patagonicus) seeking shelter in the lee of small hills during a blizzard
Gentoo penguins (Pygoscelis papua) returning over sea ice to breeding colony, Signy Island, South Orkneys, Antarctica, December
King penguins (Aptenodytes patagonicus) breeding colony with adults and juveniles, snowy mountains in background. Salisbury Plain, South Georgia. November 2018
Caribbean flamingos (Phoenicopterus ruber) line of adults standing in water, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
An adult King penguin (Aptenodytes patagonicus) amongst a creche of chicks at the breeding colony. Fortuna Bay, South Georgia Island
RF - Atlantic Puffins (Fratercula arctica) flying on to the cliff top and gathering around burrows. Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland
Brunnichs guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018
Grey-headed Flying-foxes (Pteropus poliocephalus) at a colony hang together on a branch over the Yarra River (Melbourne). ÔÇ¿ These bats form part of a colony that was established here in early 2000s
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica
King penguin (Aptenodytes patagonicus) colony. Right Whale Bay, South Georgia. September
Sociable weaver (Philetairus socius) nest in quiver tree (Aloidendron dichotomum) Quiver tree forest, Keetmanshoop, Namibia
Brown fur seal (Arctocephalus pusillus) hauled out on board walk in Cape Cross seal colony, Namibia
Walrus (Odobenus rosmarus) colony resting, Vaygach Island, Arctic, Russia, July
House sparrow (Passer domesticus) colony in building, Northern Morocco
Acrobat ant (Crematogaster sp) tending aphids on white oak (Quercus sp) Washington State Park, Pennsylvania, USA September
RF- White storks (Ciconia ciconia) nesting on poles erected by the city of Caceres in Extremadura, Spain, to compensate for the removal of an old farmhouse supporting a colony of storks
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
Snares crested penguin (Eudyptes robustus) colony standing on rocky shoreline behind band of washed up kelp, with a New Zealand fur seal (Arctocephalus forsteri) resting on rock beside them
Northern gannets (Morus bassanus) breeding colony nesting on sea cliffs, Hermaness, Shetland, Scotland, UK. July
Heermann's gull (Larus heermanni) breeding colony in flight and restless due to presence of pair of Peregrine falcon (Falco peregrinus)
Heermann's gulls (Larus heermanni) choosing and protecting nesting site. Rasa Island Special Biosphere Reserve, Sea of Cortez, Mexico. April
King penguin (Aptenodytes patagonicus) colony at dawn, Salisbury Plain, South Georgia
Blade fire coral (Millepora complanata) colonies breaking wave, protection against hurricanes. Guanaja Island, Honduras. Caribbean Sea
Diver inspecting Staghorn coral (Acropora cervicornis) and Finger coral (Porites porites) colonies. Roatan Island, Honduras. Caribbean Sea
Juvenile Southern elephant seals (Mirounga leonina) practice fighting near a King penguin (Aptenodytes patagonicus) colony, Gold Harbour, South Georgia Island
Aerial view of Caribbean flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
Royal penguin (Eudyptes schlegeli) colony, Macquarie Island, Australian Territory
Green tree ants (Oecophylla smaragdina) defending their leafy nest in a low shrub, Cooktown, Queensland, Australia
Sub-Antarctic / Brown skua (Stercorarius antarcticus) flying away with egg stolen from a Royal penguin colony (Eudyptes schlegeli), Macquarie Island, Australian Territory. December
King penguin (Aptenodytes patagonicus) chicks gathered in a creche, Fortuna Bay, South Georgia Island
Leopard seal (Hydrurga leptonyx) rests, with King penguins (Aptenodytes patagonicus) in the background. Right Whale Bay, South Georgia Island
Subantarctic or Brown skua (Stercorarius antarcticus) stealing an egg from a Royal penguin colony (Eudyptes schlegeli) on Macquarie Island, Australian Territory
King penguin (Aptenodytes patagonicus) breeding colony with chicks in creche, snow covered mountain in background. Andrews Bay, South Georgia. October 2017