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Roe deer (Capreolus capreolus) male buck in woodland, Peerdsbos, Brasschaat, Belgium August
Roe Deer (Capreolus capreolus) in woodland, Peerdsbos, Brasschaat, Belgium, June
Bengal tiger (Panthera tigris tigris), female resting and looking up Kanha National Park, India
Black-footed rock wallaby (Petrogale lateralis), Cape range National Park, Exmouth, Western Australia
North American river otter (Lontra canadensis) two in water with autumnal trees reflected in the water, Acadia National Park, Maine, USA, October
Grey seal (Halichoerus grypus) scavenging fish around trawlers in Lerwick harbour, Shetland Isles, Scotland, UK, October 2012
Tricolour merle Collie puppy, Indie, age 10 weeks
Red Squirrel (Sciurus vulgaris) male standing alert, Cairngorms National Park, Highlands, Scotland, UK, May
Zebu Cattle (Bos primigenius indicus), Los Llanos, Colombia, South America
Young rabbit hiding in grass, Okunoshima Rabbit island, Takehara, Hiroshima, Japan
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Greylag geese (Anser anser). Cley, Norfolk, UK, March. Digital composite
German Hunting Terrier, young bitch, age 9 months, sitting in straw
Masai Giraffe (Giraffa camelopardalis tippelskirchi) juveniles, Masai Mara Game Reserve, Kenya
Black Lemur (Eulemur macaco) male, Nosy Komba, Madagascar
Grey squirrel (Sciurus carolinensis) adult and baby, against white background
Northern raccoon (Procyon lotor), group standing on branch, captive
African horned viper (Cerastes cerastes), captive
Red Fox (Vulpes vulpes) curled up, with an eye open. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Female Harpy Eagle (Harpia harpyja) perched close to its nest. Pousada Currupira d Araras, south west Brazil
Himalayan marmot (Marmota himalayana) sun bathing on rock, Mount Kailash, Tibet
Northern gannet (Morus bassana) in flight, Hermaness National Nature Reserve, Shetland, Scotland, July
Wild Cat (Felis silvestris) in mist. Vosges, France, August
Arctic Fox (Alopex / Vulpes lagopus) at sunset, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Aggressive Mute swan (Cygnus olor) Upper Bavaria, Germany, Europe
On a misty morning, female Grizzly bear (Ursus arctos horribilis) standing up in alert and looking around with her cubs for danger, while feeding on sedges, Khutzeymateen Grizzly Bear Sanctuary
Northern Goshawk(Accipiter gentilis) on a branch, Bulgaria, February
RF- Great grey owl (Strix nebulosa) in flight, Finland, March
RF - Head portrait of Spotted eagle owl (Bubo africanus), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February 2016
Long-eared owl (Asio otus) perched in tree amongst autumn leaves, Czech Republic, October. Captive
Little owl (Athene noctua) perched on a fence post, stretching its wings, Castro Verde, Alentejo, Portugal, April
Bateleur Eagle (Terathopius ecaudatus) female, stretching wings Kgalagadi Transfrontier Park, South Africa. January
RF- Common frog (Rana temporaria) portrait, Cornwall, UK, January
Red Fox (Vulpes vulpes) portrait in an urban area. Glasgow, Scotland. May
Sanderling (Calidris alba) running along the tide-line, reflected in the mud, Norfolk, England, UK. December
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
RF- Peregrine Falcon (Falco peregrinus) landing on moorland with its wings spread. Scotland, UK, March
Great Tit (Parus major) perched on branch with yellow from autumn trees in the background. Southern Norway, October
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April
Big-legged Bug or Squash Bug (Coreidae) on Heliconia Flower. Yasuni National Park, Amazon Rainforest, Ecuador. South America
RF- Hyacinth macaw (Anodorhynchus hyacinthinus) in its nest hole. Pousada Aguape Lodge, Mato Grosso do Sul State, Brazil. September
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
Erect-crested penguins (Eudyptes sclateri) pair on coast, Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Bengal Tigers (Panthera tigris tigris) adult female drinking water, her juvenile cub beside her. Bandhavgarh National Park, India, April
Bengal Tiger (Panthera tigris tigris) captive, with leaves digitally added
RF- Tiger (Panthera tigris) 14-month Lakshmi cub resting in tree, Bandhavgarh National Park, India. Endangered species
Female Cheetah (Acinonyx jubatus) standing in long grass on the savannah, Okavango Delta, Botswana, Africa
American pine marten (Martes americana) standing on hind legs in deep snow, Yellowstone National Park, USA. January