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Northern / Hen Harrier (Circus cyaneus) and Kestrel (Falco tinnunculus) below, fighting in flight. Vosges, France, February
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Serotine bat (Eptesicus serotinus) showing teeth, Captive, UK
Silverback male Eastern lowland gorilla (Gorilla beringei graueri) intimidation display by chest beating, equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Arctic hares (Lepus arcticus) fighting, Northeast Greenland National Park, Greenland. Veolia Environnement Wildlife Photographer of the Year 2009: Mammal Behaviour category: Runner-up
Wild Horses / mustangs, two stallions rearing up fighting, Pryor Mountains, Montana, USA, June
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
King cobra (Ophiophagus hannah) in strike pose, Malaysia
European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July
Frill-necked lizard (Chlamydosaurus kingii), with collar spread in threat display. Alice Springs, Northern territory, Australia
Frill-neck Lizard (Chlamydosaurus kingii), displays on a termite mound. Northern Territory, Australia
Fosa / Fossa (Cryptoprocta ferox) male snarling, dry deciduous forest, Kirindy Forest, Western Madagascar, IUCN vulnerable species
Tokay gecko (Gekko gecko) enacting a defensive display towards a perceived threat, Phuket Island, Thailand, March. Controlled conditions
Yellow spotted monitor standing threat display {Varanus panoptes panoptes} Queensland, Australia
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Crested caracaras (Caracara cheriway), midair aggressive interaction between two for access to perch, Martin Refuge, Rio Grande Valley, Texas, USA, March
Red-headed woodpeckers (Melanerpes erythrocephalus), pair fighting with European Starling (Sturnus vulgaris) that is trying to take over the woodpeckers nest hole, Montezuma National Wildlife Refuge
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Kestrel (Falco tinnunculus) fighting Hungary, April
Great white shark (Carcharodon carcharias) breaking surface with mouth open. Guadalupe Island, Mexico
Stag beetle (Lucanus cervus) males fighting on oak tree branch, Elbe, Germany, June
African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana
RF - Polar Bears (Ursus maritimus) males fighting, Churchill, Canada, November
Camargue horse, two, one with mouth open. Bouches du Rhone, France May
Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August
Black bears (Ursus americanus) standing on back legs, fighting, Minnesota, USA, June
Northern goshawk (Accipiter gentilis) flying with squirrel prey, Finland. March
King cobra (Ophiophagus hannah) in strike pose with mouth open, tongue out and glottis (hole like structure in mouth) clearly visible. Malaysia
Bombardier Beetle (Pheropsophus jessoensis) protecting itself by ejecting a boiling, noxious chemical spray, Nagasaki, Japan
Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October
Avocet (Recurvirostra avosetta) trying to drive Shelduck (Tadorna tadorna) pair away from breeding grounds, Oare marshes, RSPB Greater Thames Futurescapes Project, North Kent, UK, June
Egyptian cobra (Naja haje) with head raised up and hood expanded, near Ouarzarte, Morocco
Rufous-tailed hummingbird (Amazilia tzacatl) territorial fighting around Heliconia flower (Heliconia latispatha) La Selva, Costa Rica
Two Jaguars (Panthera onca) snarling at each other on the riverbank, Pantanal, Brazil. September
Two Goldfinches (Carduelis carduelis) squabbling over Common teasel (Dipsacus fullonum) seeds in winter, Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. 2020VISION Exhibition
Emma Grays Forest Lizard (Calotes emma) in threat display, Xishuangbanna National Nature Reserve, Yunnan Province, China, March
Wild Boar (Sus scrofa) males fighting in snow. The Netherlands, January
Grevy zebra (Equus grevyi) stallions fighting, Samburu National Reserve, Kenya
African elephant {Loxodonta africana} charging, Etosha national park, Namibia
