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Little owl {Athene noctua) pair perched, courtship behaviour, Spain
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Masai giraffe {Giraffa camelopardalis} mother nuzzling baby, Lower Masai Mara GR, Kenya
Palomino Peruvian paso mare and foal, Sante Fe, New Mexico, USA
Plains zebra (Equus quagga) grooming foal, Kruger National Park, South Africa, July
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Tabby kitten, Picasso, 9 weeks, head to head with Yellow labrador puppy, 8 weeks
Female Orang Utan (Pongo pygmaeus) [Sandy, born 29.04.82] sitting, holding two young. One [Samboja, born 09.06.05] is her own offspring, the other [Dayang
Adult Mountain gorilla (Gorilla beringei beringei) holding baby, Hirwa group, Volcanoes National Park, Rwanda
Sea Otter floating at sea surface holding pup. Alaska (Enhydra lutris) USA
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
European Bee-eater (Merops apiaster) pair, male passing Dragonfly prey to female, Pusztaszer, Hungary, May 2008
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Baby rabbit with a Roborovski Hamster (Phodopus roborovskii) sitting on its head
Arctic wolf (Canis lupus) male and female, captive
Eurasian lynx (Lynx lynx) kitten, aged eight months, nuzzling its mother, breeding and reintroduction program, Germany. Captive. March
Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. June
Young orphan Elephants (Loxodonta africana) kissing. David Sheldrick Wildlife Trust Nairobi Elephant Nursery, Kenya, July 2010
Grey wolf (Canis lupus), adult with cub, captive, USA
Hoffmanns Two-toed sloth (Choloepus hoffmanni) mother and baby, aged 2 months, in tree, Costa Rica. Rescued and released by Aviarios Sloth Sanctuary
Jaguar (Panthera onca) mother grooming with four month old cub, native to Southern and Central America, captive
Tabby kittens, Stanley and Fosset, 12 weeks old, walking together
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
Cute red and red-and-white Cavapoo puppies, 5 weeks, hugging, against white background
Spotted owlet (Athene brama) pair on a branch, Bharatpur / Keoladeo Ghana National Park, India
Hyacinth macaws (Anodorhynchus hyacinthinus) mutual preening, Estrada Parque, Pantanal, Brazil
Albino Common Iguana (Iguana iguana) captive pair from Tom Crutchfields collection, from Caribbean and South and Central America
RF - Scarlet macaw (Ara macao) pair preening, Osa Peninsula, Costa Rica
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
California sea lions (Zalophus californianus) two resting with flippers round each other, Monterey Bay, California, USA, September
Harvest mice (Micromys minutus) on teasel seed head. Dorset, UK, August. Captive
Young Japanese macaque (Macaca fuscata) kissing another macque in the thermal hot springs, Jigokudani, Japan, January
European rabbit (Oryctolagus cuniculus) two young rabbits, or kittens, touch noses, Norfolk, UK, June. Non-ex
Mute swan (Cygnus olor) pair courting. Walthamstow reservoir, London, UK
Indian fox (Vulpes bengalensis) pups at play by a den in the Little Rann, Kutch, Gujarat, India. Honorable mention in the Land Mammals category of the BigPicture Competition 2014
Grizzly bear (Ursus arctos) female and nuzzling cub, Lake Clark National Park, Silver Salmon, Alaska. July
Two Exmoor ponies (Equus ferus caballus), semi-feral native breed, rubbing noses, Exmoor National Park, Somerset / Devon, England. November
Female Snow leopard (Panthera uncia) with her three month cub looking up at its mother, BioParc Doue la Fontaine, France. Captive
Eurasian lynx (Lynx lynx) kitten, age six weeks, showing affection towards its mother. Breeding and reintroduction program, Germany. Captive. July
Mother and three months old Snow leopard (Panthera uncia) cub cleaning each other, Bioparc of Doue La Fontaine, France. September
Eurasian lynx (Lynx lynx) grooming its kitten, aged eight months, breeding and reintroduction program, Germany. Captive. March
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Greater flamingo (Phoenicopterus roseus) pair at rest in water, Cape Town, South Africa, July
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
African bush elephant, (Loxodonta africana) two head to head playing, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Erect-crested penguins (Eudyptes sclateri) feeding young chick. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Welsh Ponies (Equus caballus) engaged in mutual grooming, Llanbedr Hill, Powys, Wales, August
