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Quaking aspen trees (Populus tremuloides) and conifers with dusting of snow, Dome Hill above Dawson City, Yukon Territories, Canada, September 2013
Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya
Detail of the pectoral fin of Roundhead parrotfish (Scarus viridifucatus) coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Pig-nosed frog 1+Nasikabatrachus sahyadrensis+2 in leaf litter, Western Ghats, India
Detail of melting ice on Jokulsarlon beach. Southern Iceland, Europe, November 2012
Leafless tree canopy silhouetted in winter. Gannochy, Scotland, February
Common poppy (Papaver rhoeas) bud, Polly / Porth Joke, Pentire, Newquay, Cornwall, UK, July
Ice fragment on crack in Lake Baikal ice, Siberia, Russia, March
Map lichen (Rhizocarpon geographicum) growing on schist boulder. Nordtirol, Austrian Alps, 2250 metres above sea level. August
Indian moon / Indian luna Moth (Actias selene) close up of defensive spines on caterpillar. Captive
Carrion Crow (Corvus corone) encircled by flock of Starlings (Sturnus vulgaris) resting on the beach in Blackpool, England, October
Slender sapsucking slug (Thuridilla gracilis) on surface of Royal seasquirt (Polycarpa aurata) Aljui Bay, Waigeo Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Jackdaws (Corvus monedula) and Rooks (Corvus frugilegus) mixed winter flock taking flight from a frost covered field of crop stubble, Derbyshire, UK, January (non-ex)
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
RF- Chanterelle mushroom (Cantharellus cibarius), gill detail
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean
Light Bulb Sea Squirt (Clavelina lepadiformis). Channel Islands, UK, June
Detail of palm leaf, backlit in sunshine, New Caledonia
Great yellow gentian (Gentiana lutea) leaves. Haute-Savoie, Massif des Bauges, French Alps, May
Brain coral (Diploria strigosa) photographed under blue light to excite the fluorescence, Little Cayman, Cayman Islands, British West Indies, Caribbean Sea
Abstract reflection of reeds in rippled water, Westhay Moor SWT reserve, Somerset Levels, Somerset, England, UK, June 2011
Palm Frond detail. Mahahual Penninsula, South Yucatan Peninsula, Mexico
RF - Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
Close up of pattern of adult Emperor angelfish (Pomacanthus imperator) Tulamben, Bali, Indonesia. Java Sea
Detail of sponge (Chalinula nematifera) Maldives, Indian Ocean
Broken ice abstract, Jokulsarlon glacier lagoon. Southern Iceland, Iceland, Europe, March 2011
Detail in frozen waterfall. Selijalandfoss waterfall, Southern Iceland, Europe, March 2011
Beech (Fagus) leaves against white background. Scotland
Various pollen grains from several species of UK plant. False coloured scanning electron micrograph. Digital composite
Aerial view saltmarshes of Bahia de Cadiz Natural Park, Huelva, Andalucia, Spain, March 2008
Aerial view of Grand Prismatic thermal spring in Yellowstone National Park, USA. The bright colours are natural, coming from thermophile bacteria in the water
Abstract of reeds in front of lake with reflections, Plitvice Lakes National Park, Lika, Croatia, Europe, October 2011
Leaves floating on water with colourful reflections, Plitvice Lakes National Park, Lika, Croatia, Europe, October 2011
Sunflower (Helianthus annuus) Spain
Rothschilds giraffe (Giraffa camelopardalis rothschildi) close up of young calf skin pattern, captive
Grevy's zebra (Equus grevyi) close up, captive
Cotton Grass (Eriophorum vaginatum) growing in water. Goldenstater Moor, Niedersachsen, Germany
Looking up through the leaves of Wild sarsaparilla (Aralia nudicaulis) in woodland, Cape Cod, Massachusetts, USA, May
Aerial view of the rivers and saltmarshes of the Bahia / Bay de Cadiz Natural Park, Andalucia, Spain, March 2008
River patterns from melting glacier. Olfusa river, Southwest Iceland, July 2009
Aerial view of Lake Natron, Rift Valley, Tanzania, Africa, August 2009
Detail in ice pattern formations, the Netherlands
Detail of leaf of Cabbage Palmetto (Sabal palmetto) Sanibel Island, Florida, USA
Silhouette of a Green Sea Turtle (Chelonia mydas) passes overhead. West Maui, Hawaii